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Kays Roland - Mammals of North America

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Kays Roland Mammals of North America
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    Mammals of North America
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Mammals of North America illustrates all 462 known mammal species in the United States and Canada--each in beautiful color and accurate detail. With a more up-to-date species list than any other guide, improved facing-page descriptions, easier-to-read distribution maps, updated common and scientific names, and track and scat illustrations, this slim, light, and easy-to-use volume is the must-have source for identifying North American mammals.;Artists credits -- About the Author -- Acknowledgments -- Introduction -- Species Plates -- Plate 1. Opossum and Armadillo -- Plate 2. Eastern Sorex -- Plate 3. Northern Shrews -- Plate 4. Aquatic Shrews and Skulls -- Plate 5. Inland Sorex with Skulls -- Plate 6. Coastal Sorex with Skulls -- Plate 7. Other Shrews -- Plate 8. Western Moles -- Plate 9. Eastern Moles -- Plate 10. Pikas -- Plate 11. Western Cottontails -- Plate 12. Pygmy Rabbits and Short-eared Cottontails -- Plate 13. Eastern Cottontails -- Plate 14. Northern Hares -- Plate 15. Jackrabbits -- Plate 16. Porcupine and Sewellel -- Plate 17. Large Aquatic Rodents -- Plate 18. Marmots -- Plate 19. Eastern and Tropical Tree Squirrels -- Plate 20. Western Tree Squirrels -- Plate 21. Red and Flying Squirrels -- Plate 22. Prairie Dogs -- Plate 23. Large, Spotted Ground Squirrels -- Plate 24. Bushy-tailed, Flecked Ground Squirrels -- Plate 25. Thin-tailed, Unmarked Ground Squirrels -- Plate 26. Small, Spotted Ground Squirrels -- Plate 27. Stripy Ground Squirrels -- Plate 28. Eastern and Rocky Mountain Chipmunks -- Plate 29. Southwest Chipmunks -- Plate 30. Southern California Chipmunks -- Plate 31. Northwestern Coastal Chipmunks -- Plate 32. Northwestern Inland Chipmunks -- Plate 33. Northern Thomomys -- Plate 34. Cratogeomys and Southern Thomomys -- Plate 35. Geomys -- Plate 36. Western Desert Perognathus -- Plate 37. Great Plains Perognathus -- Plate 38. Liomys and Spiny Chaetodipus -- Plate 39. Smooth Chaetodipus -- Plate 40. Kangaroo Mice and Small Four-toed Kangaroo Rats -- Plate 41. Large Four-toed Kangaroo Rats -- Plate 42. Mid-California Five-toed Kangaroo Rats -- Plate 43. Southern California Five-toed Kangaroo Rats -- Plate 44. Other Five-toed Kangaroo Rats -- Plate 45. Jumping Mice -- Plate 46. Eastern and Central Woodrats -- Plate 47. Furry-tailed Woodrats -- Plate 48. Woodrats and Rattus -- Plate 49. Grasshopper Mice and Oryzomys -- Plate 50. Central and Eastern Tiny Mice -- Plate 51. House and Western Tiny Mice -- Plate 52. Northern Deermice -- Plate 53. Southeastern Peromyscus and Relatives.;Plate 54. Far West Peromyscus -- Plate 55. Peromyscus truei and boylii Groups -- Plate 56. Sigmodon -- Plate 57. Arborimus Voles -- Plate 58. Red-backed Voles -- Plate 59. Coastal Microtus -- Plate 60. Other Western Voles -- Plate 61. Northern Voles -- Plate 62. Typical Voles -- Plate 63. Eastern Voles and Bog Lemmings -- Plate 64. Northern Lemmings -- Plate 65. Ghost and Leaf-nosed Bats -- Plate 66. Molossid Bats -- Plate 67. Lasiurine Bats -- Plate 68. Giant-eared Bats -- Plate 69. Longer-eared Myotis -- Plate 70. Western Myotis 1 -- Plate 71. Western Myotis 2 -- Plate 72. Eastern Myotis -- Plate 73. Other Eastern Bats -- Plate 76. Temperate Cats -- Plate 76. Temperate Cats -- Plate 77. Tropical Cats -- Plate 78. Canis -- Plate 79. Smaller Foxes -- Plate 80. Larger Foxes -- Plate 81. Bears -- Plate 82. Procyonids -- Plate 83. Aquatic Mustelids -- Plate 84. Big Mustelids -- Plate 85. Mustela -- Plate 86. Spotted and Hog-nosed Skunks -- Plate 87. Mephitis -- Plate 88. Otariid Seals -- Plate 89. Phocid Seals -- Plate 90. Grayish Seals -- Plate 91. Big Seals and Manatee -- Plate 92. Antelopes and Pigs -- Plate 93. Goats and Sheep -- Plate 94. Bison and Nilgai -- Plate 95. Arctic Ungulates -- Plate 96. Elk and Moose -- Plate 97. Native Deer -- Plate 98. Exotic Ungulates -- Plate 99. Large Whales without Dorsal Fins -- Plate 100. Enormous Whales with Dorsal Fins -- Plate 101. Large Whales with Dorsal Fins -- Plate 102. Small White and Gray Whales -- Plate 103. Black Dolphins -- Plate 104. Unstriped, Beaked Dolphins -- Plate 105. Striped, Beaked Dolphins -- Plate 106. Striped, Blunt-nosed Dolphins and Porpoises -- Plates 107. and 108. Bow-riding Dolphins and Whales -- Plate 109. Beaked Whales -- Plate 110. Heads of Male Mesoplodon -- Plates 111. and 112. Whale and Dolphin Dive Sequences (not to scale) -- Plate 113. Carnivore Scats -- Plate 114. Vegetarian Scats -- Glossary -- Appendix.

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P RINCETON F IELD G UIDES
For additional titles in this series visit www.guides.princeton.edu

Albatrosses, Petrels, and Shearwaters of the World, Derek Onley and Paul Scofield

Birds of Australia, Seventh Edition, Ken Simpson and Nicolas Day

Birds of Chile, Alvaro Jaramillo

Birds of East Africa Kenya, Tanzania, Uganda, Rwanda, and Burundi, Terry Stevenson and John Fanshawe

Birds of India Pakistan, Nepal, Bangladesh, Bhutan, Sri Lanka, and the Maldives, Richard, Grimmett, Carol Inskipp, and Tim Inskipp

Birds of Northern India, Richard Grimmett and Tim Inskipp

Birds of Peru, Thomas S. Schulenberg, Douglas F. Stotz, Daniel F. Lane, John P. ONeill, and Theodore A. Parker III

Birds of Southeast Asia, Craig Robson

Birds of Southern Africa, Ian Sinclair, Phil Hockey, and Warwick Tarboton

Birds of Thailand, Craig Robson

Birds of the Middle East, R. F. Porter, S. Christensen, and P. Schiermacker-Hansen

Birds of the West Indies, Herbert Raffaele, James Wiley, Orlando Garrido, Allan Keith, and Janis Raffaele

Birds of Western Africa, Nik Borrow and Ron Demey

Caterpillars of Eastern North America A Guide to Identification and Natural History, David L. Wagner

Mammals of Europe, David W. Macdonald and Priscilla Barrett

Mammals of North America, Second Edition, Roland W. Kays and Don E. Wilson

Marine Mammals of the North Atlantic, Carl Christian Kinze

Minerals of the World, Ole Johnsen

Nests, Eggs, and Nestlings of North American Birds Second Edition, Paul J. Baicich and Colin J.O. Harrison

Raptors of the World, James Ferguson-Lees and David A. Christie

Reptiles and Amphibians of Europe, E. Nicholas Arnold

Sharks of the World, Leonard Compagno, Marc Dando, and Sarah Fowler

Stars and Planets: The Most Complete Guide to the Stars, Planets, Galaxies, and the Solar System Fully Revised and Expanded Edition, Ian Ridpath and Wil Tirion

Whales, Dolphins, and Other Marine Mammals of the World, Hadoram Shirihai and Brett Jarrett

Mammals of
North America

Second Edition

ROLAND W. KAYS and DON E. WILSON

Princeton University Press Princeton and Oxford A RTISTS CREDITS Sandra - photo 1

Princeton University Press

Princeton and Oxford

A RTISTS CREDITS
Sandra Doyle/Wildlife Art Ltd.: Plates 88-91, 99-112
Nancy Halliday: Plates 2-9, 28-32
Elizabeth McClelland: Plates 10-18, 33-49, 92-98
Consie Powell: Plates 76-87
Wendy Smith: Plates 50-55, 65-75
Todd Zalewski: Plates 1, 19-27, 56-64
Cover illustration: Elizabeth McClelland
Scat illustrations: Diane Gibbons
Track illustrations: Susan C. Morse and Jesse Guertin Copyright 2009 by Princeton University Press
Published by Princeton University Press
41 William Street
Princeton
New Jersey 08540

In the United Kingdom:
Princeton University Press
6 Oxford Street Woodstock
Oxfordshire OX20 1TW

All Rights Reserved
Library of Congress Cataloging-in-Publication Data Kays, Roland, 1971
Mammals of North America/Roland W. Kays and Don E. Wilson.2nd ed.
p. cm.(Princeton field guides)
Includes bibliographical references and index.
ISBN 978-0-691-14278-4 (cloth : alk. paper)
ISBN 978-0-691-14092-6 (pbk : alk. paper)
1. MammalsNorth AmericaIdentification. I. Wilson, Don E. II. Title.
QL715 .K38 2009
599.097dc22 2009001417

British Library Cataloging-in-Publication Data is available This book has been composed in Galliard (main text) and ITC Franklin Gothic (headings and tabular material) Printed on acid-free paper.

press.nathist.edu

Edited and designed by D & N Publishing, Baydon, Wiltshire, UK

Printed in Italy by Eurografica SPA

10 9 8 7 6 5 4 3 2 1

Roland W. Kays is the Curator of Mammals at the New York State Museum. His research centers on the ecology and conservation of temperate and tropical mammals, especially carnivores.

Don E. Wilson is Chairman of the Department of Vertebrate Zoology at the Smithsonian Institutions National Museum of Natural History. Recipient of a Smithsonian Institution Award for Excellence in Tropical Biology and a U.S. Fish and Wildlife Service Outstanding Publication Award, he is the author or coauthor of more than 225 scientific papers and 20 books, including three volumes on bats, The Smithsonian Book of North American Mammals, Mammal Species of the World, and Handbook of Mammals of the World.

C ONTENTS
ACKNOWLEDGMENTS

The trick to writing a good field guide is to collect the scattered information on species identification and concentrate it into one concise volume. As a whole, North American mammals are a well-studied group, and the details in this book are largely a credit to the work produced by generations of mammalogists. Although books and journal articles provided some of this detail, no field guide could be completed without substantial research in natural history museums, and this guide is no exception. Our own institutions, the New York State Museum and the National Museum of Natural History, provided the collections, library facilities, and a wide variety of witting and unwitting ancillary support. In addition, we thank the Denver Museum of Nature and Science, the University of Colorado Museum, and the Field Museum, for access to their collections. A specimen in the drawer is much easier to bring to life when an expert helps point out the relevant characters. For this museum and identification help we thank:

Greg Anderson, Andrea Bixler, Joe Bopp, Harold Broadbooks, Mike Carleton, Chris Conroy, Linda Gordon, Lawrence R. Heaney, Al Hicks, Rosanne Humphrey, Cheri A. Jones, Zack Knight, Bill Longland, Chris Maser, Jesus Maldonado, James Mead, Bruce Patterson, Jim Patton, Toni Piaggio, John Phelps, Roger A. Powell, Eric Rickart, Dave Schmidt, Michi Schulenberg, Andrew T. Smith, Bob Smith, William Stanley, and Westarp Wissenschaften. Additionally, we thank William Gannon for his assistance with chipmunk vocalizations, Dan Simberloff for his help with introduced species, and Al Hicks for plunging into caves to help us get the bats just right. Tim Page carefully read and corrected the entire text. A special thanks to Wade Sherbrooke and the staff at the Southwest Research Station for their hospitality during one of our field trips.

The range maps were provided in a GIS for this field guide by Wes Sechrest. In collaboration with many conservation, museum, and academic groups including Conservation International, the Institute of Applied Ecology, and the International Union for the Conservation of Nature (IUCN), Sechrest has compiled distribution information for all mammal species in the world. The initial products, extent of occurrence maps for all mammal species, will soon be freely available on the new Species Information Service (SIS) of the IUCN.

A field guide is part science and part art. For help and critique of artwork we thank:

Catherine Chapman, Marlene Hill Donnelly, Ben Flemer, Patricia Kernan, Clara Richardson Simpson, David Steadman, and Karen Teramura, the staff at WILDlifeART, and Christa Wurm. Thanks to Keeping Tracks (www.keepingtrackinc.org) founder and Program Director Susan Morse for taking the time off their citizen-based wildlife monitoring, educational, and conservation programs to provide her expertise regarding the animal tracks and scat illustrations.

We thank Greg Anderson, Tom Brooks, Chris Byrne, Chip Foster, Fritz Hertel, Barret Klein, Darrin Lunde, Chris Skelton, and John Young for insightful discussions about field guide design. Thanks to Ron Gill, Adam Fox, and Dimitri Karetnikov for computer help. Our editors at Princeton University Press, Sam Elworthy and Robert Kirk, gently pushed and pulled at appropriate points throughout the process, contributing greatly to the timely completion of the book.

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