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Keeley Sterling C. - Introduction to California Chaparral

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Keeley Sterling C. Introduction to California Chaparral
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ABOUT THE AUTHORS

Sterling C Keeley is Professor of Botany at the University of Hawaii in - photo 1

Sterling C. Keeley is Professor of Botany at the University of Hawaii in Honolulu. She served as the scientific adviser for the Chaparral Hall at the Los Angeles County Museum of Natural History and edited The California Chaparral: Paradigms Reexamined (1989). In California, her research focused on the fire-following plants of the chaparral.

Ronald D Quinn is Professor of Biological Sciences at California State - photo 2

Ronald D. Quinn is Professor of Biological Sciences at California State Polytechnic University, Pomona. He has published widely on effects of fire and herbivory on chaparral, and other mediterranean ecosystems of the world.

Marianne D. Wallace has been a natural science illustrator and educator for over 30 years. She lives in the foothills of southern California, sharing her toyon-bordered backyard with wood rats, Mule Deer, Wrentits, and other chaparral species.

ACKNOWLEDGMENTS

I give heartfelt thanks to my spouse, Barbara Ellis-Quinn, who in countless ways has acted as a patient and supportive midwife for the rather protracted birth of this book. She reviewed the manuscript in its entirety and created a quiet office and a warm home for writing. My friend Ken Montgomery, a botanist and horticulturist with a true love of plants native to California, provided hours of discussion and dreaming when I first thought to write this book a half lifetime ago. He has given me support, encouragement, and suggestions when they were most needed. The Rancho Santa Ana Botanic Garden of Claremont, California, has provided many days of pleasure and inspiration, as well as a place to take photographs for this book. Steve Shirley and his airplane gave me a new perspective and some photographs of wildfires at the urban-wildland interface. Rick Halsey's boundless enthusiasm was inspirational, as was his generosity with his ideas. Mike Raugh, mathematician and accomplished naturalist, reviewed a large part of the manuscript. Colleagues Chris Brady, Glenn Stewart, and Gary Wallace reviewed various parts of the manuscript. William D. Wagner provided insights on birds. Jim Dole, who first introduced me to the delights and challenges of chaparral ecology, set me on a path that has lasted a lifetime. Finally, I am grateful to students of chaparral biology at California State Polytechnic University, Pomona, who over the years have been an endless source of enthusiasm to me for all things to do with chaparral.

Ronald D. Quinn

February 2006


This book would never have come into being were it not for the inspiration and support of many people. For starting my career in chaparral research and encouraging my efforts along the way, Harold Mooney of Stanford University, deserves my deepest gratitude. Nancy Coile, of the University of Georgia, provided the initial inspiration to write this book while visiting and collecting ceanothus. My sister, Christine Thompson, willingly took hikes with me while I photographed and pontificated, read countless drafts, and buoyed up my enthusiasm when it flagged. Frank Hovore spent many hours providing interesting information about insects and animals of the chaparral, and reading drafts, as well as providing some fabulous photos. Steve Davis, Robert Gustafson, Linda Hardie-Scott, Charles Hogue, Dave Minor, Philip Rundel, Timothy Thomas, Sherry Wood, and Paul Zedler provided wonderful photographs and have been waiting for them to be returned since the early 1990s. Their generosity and patience are much appreciated. Shirleen Gudmuson, Christa Hatch, Linda Kate Schroeder, and Peter Vroom all contributed to editing, reviewing, and refining the manuscript during its various incarnations. My students at Whittier College allowed me to share my delight in the chaparral and to do original research that would not have been possible without them. They taught me to see the natural world in new ways, for which I am truly grateful. Thanks to Phil Rundel and one anonymous reviewer for contributing to the quality of the book. And finally, our thanks to the editors, Phyllis Faber, who suggested we work together on this project, and Doris Kretschmer, who worked out the details and kept us on track.

Sterling C. Keeley

February 2006

ART CREDITS

Photographs were generously provided by a number of colleagues. Individual slide credits are given below; those not listed are from the authors' personal collections.

Plates

STEPHEN DAVIS, Pepperdine University

JACQUES DESCLOITRES, courtesy of NASA

WILLIAM FOLLETTE

ROBERT GUSTAFSON, Los Angeles Country Museum of Natural History

LINDA HARDIE-SCOTT, The Nature Conservancy

CHARLES HOGUE, Los Angeles County Museum of Natural History

FRANK HOVORE/DEDE GILMAN, Hovore Associates

GLENN KEATOR

PHILIP RUNDEL, University of California at Los Angeles

TIMOTHY THOMAS, U.S. Fish and Wildlife Service

SHERRY WOOD, Gonzaga University

PAUL ZEDLER, University of Wisconsin

Figures

Chamise shrubs drawn after Arthur W. Sampson, Plant succession on burned chaparral lands in northern California, Calif. Agric. Exper. Stn. Bull. 685, 1944.

Drawn in part from Hazel Gordon and Thomas C. White, Ecological Guide to Southern California Chaparral Plant Series: Transverse and Peninsular Ranges: Angeles, Cleveland, and San Bernardino National Forests, RS-ECOL-TP-005 (San Francisco: Cleveland National Forest, U.S.D.A. Forest Service, 1994).

Cone redrawn with permission from Jim W. Dole and Betty B. Rose, Shrubs and Trees of the Southern California Coastal Region (North Hills, Calif.: Foot-loose Press, 1996).

Whiptail drawn after photograph by Nathan W. Cohen in Alden H. Miller and Robert C. Stebbins, The Lives of Desert Animals in Joshua Tree National Monument (Berkeley and Los Angeles: University of California Press, 1964).

of Klaus W. H. Radtke, Living More Safely in the Chaparral-Urban Interface, General Technical Report PSW-67 (Berkeley, Calif.: Pacific Southwest Forest and Range Experiment Station, U.S. Department of Agriculture, 1983).

CHAPTER 1
THE CALIFORNIA CHAPARRAL

CHAPARRAL IS BOTH a vegetation type and the name given to the community of - photo 3

CHAPARRAL IS BOTH a vegetation type and the name given to the community of - photo 4

CHAPARRAL IS BOTH a vegetation type and the name given to the community of coadapted plants and animals found in the foothills and mountains throughout California. The chaparral vegetation is composed of a diverse assemblage of different species of evergreen drought- and fire-hardy shrubs. Seen from the car window or scenic lookout, chaparral looks like a soft bluish green blanket gently covering the hills. Up close, however, this blanket no longer appears soft. Instead, what is revealed is a nearly impenetrable thicket of shrubs with intertwined branches and twigs with hard leaves and stiff and unyielding stems. The shrubs are well adapted to the rigors of long, hot, dry summers and unpredictable winter rainfall that are characteristic of California's mediterranean climate. Chaparral is especially extensive in the central and southern parts of the state, but it covers large areas of northern California as well ().

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