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Thad Q. Bartlett - The Gibbons of Khao Yai

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PRIMATE FIELD STUDIES
Series Editors:
Robert W. Sussman, Washington UniversitySt. Louis
Natalie Vasey, Portland State University
Series Editorial Board:
Simon Bearder, Oxford-Brookes University
Marina Cords, Columbia University
Agustin Fuentes, Notre Dame University
Paul Garber, University of Illinois
Annie Gautier-Hion, Station Biologique de Paimpont
Joanna Lambert, University of Wisconsin
Robert D. Martin, Field Museum
Deborah Overdorff, University of Texas
Jane Phillips-Conroy, Washington University
Karen Strier, University of Wisconsin
Series Titles:
The Spectral Tarsier
Sharon L. Gursky, Texas A&M University
Strategies of Sex and Survival in Hamadryas Baboons: Through a Female Lens
Larissa Swedell, Queens College, The City University of New York
The Behavioral Ecology of Callimicos and Tamarins in Northwestern Bolivia
Leila M. Porter, The University of Washington
The Socioecology of Adult Female Patas Monkeys and Vervets
Jill D. Pruetz, Iowa State University
Apes of the Impenetrable Forest: The Behavioral Ecology of Sympatric Chimpanzees and Gorillas
Craig B. Stanford, University of Southern California
A Natural History of the Brown Mouse Lemur
Sylvia Atsalis, Brookfield Zoo
The Gibbons of Khao Yai: Seasonal Variation in Behavior and Ecology
Thad Q. Bartlett, The University of Texas at San Antonio
PRIMATE FIELD STUDIES
Many of us who conduct field studies on wild primates have witnessed a decline in the venues available to publish monographic treatments of our work. As researchers, we have few choices other than to publish short technical articles on discrete aspects of our work in professional journals. Also in vogue are popular expositions, often written by nonscientists. To counter this trend, we have begun this series. Primate Field Studies is a venue both for publishing the full complement of findings of long-term studies and for making our work accessible to a wider readership. Interested readers need not wait for atomized parts of long-term studies to be published in widely scattered journals; students need not navigate the technical literature to bring together a body of scholarship better served by being offered as a cohesive whole. We are interested in developing monographs based on single- or multi-species studies. If you wish to develop a monograph, we encourage you to contact one of the series editors.
About the Editors:
Robert W. Sussman (Ph.D. Duke University) is currently Professor of Anthropology and Environmental Science at Washington University, St. Louis, Missouri, and past Editor-in-Chief of American Anthropologist, the flagship journal of the American Anthropological Association. His research focuses on the ecology, behavior, evolution, and conservation of nonhuman and human primates, and he has worked in Costa Rica, Guyana, Panama, Madagascar, and Mauritius. He is the author of numerous scientific publications, including Biological Basis of Human Behavior, Prentice Hall (1999), Primate Ecology and Social Structure (two volumes), Pearson Custom Publishing (2003), and The Origin and Nature of Sociality, Aldine de Gruyter (2004).
Natalie Vasey (Ph.D. Washington University) is currently Assistant Professor of Anthropology at Portland State University in Portland, Oregon. Her work explores the behavioral ecology, life history adaptations, and evolution of primates, with a focus on the endangered and recently extinct primates of Madagascar. She has presented her research at international venues and published in leading scientific journals. She is dedicated to educating students and the public-at-large about the lifestyles and conservation status of our closest relatives in the Animal Kingdom.

The Gibbons
of Khao Yai

The Gibbons
of Khao Yai
Seasonal Variation in
Behavior and Ecology
Thad Q. Bartlett
The University of Texas at San Antonio
First published 2009 by Pearson Education Inc Published 2016 by Routledge 2 - photo 1
First published 2009 by Pearson Education, Inc.
Published 2016 by Routledge
2 Park Square, Milton Park, Abingdon, Oxon OX14 4RN
711 Third Avenue, New York, NY 10017, USA
Routledge is an imprint of the Taylor & Francis Group, an informa business
Copyright 2009 Taylor & Francis. All rights reserved.
All rights reserved. No part of this book may be reprinted or reproduced or utilised in any form or by any electronic, mechanical, or other means, now known or hereafter invented, including photocopying and recording, or in any information storage or retri eval system, without permission in writing from the publishers.
Notice:
Product or corporate names may be trademarks or registered trademarks, and are used only for identification and explanation without intent to infringe.
Credits and acknowledgments borrowed from other sources and reproduced, with permission, in this textbook appear on appropriate page within text.
This book was set in 10/12 Palatino.
ISBN: 9780131915046 (pbk)
Cover Design: Margaret Kenselaar
Library of Congress Cataloging-in-Publication Data
Bartlett, Thad Q.
The gibbons of Khao Yai: seasonal variation in behavior and ecology / Thad Q. Bartlett.
p. cm. (Primate field studies)
Includes bibliographical references and index.
ISBN 0-13-191504-5
1. GibbonsBehaviorThailandUtthayan hfng Chat Khao Yai. 2. GibbonsEcologyThailandUtthayan hfng Chat Khao Yai. I. Title.
QL737.P96B355 2009
599.88209593dc22 2008025788
Dedicated to my mother, Bertrice Bartlett

Contents



It has now been many years since I first visited Khao Yai National Park to begin my dissertation research, nevertheless, I can still remember the moment when I saw wild gibbons in person for the first time. I had come to the park with Warren Brockelman, who by that time had been working in Khao Yai, off and on, for well over a decade. I was following Warren through the forest as he oriented me to the trail system and topography of the study area. We had been walking for a few hours without a glimpse of gibbons, long enough that I had already started to wonder if I would have to wait another day, or longer, to finally see the animals I proposed to study for the coming year. My anxiety was temporarily eased as we crossed over a small ravine, and Warren pointed up into the crown of an unimaginably large tree. It was a sprawling Ficus that soared over a hundred feet in the air, and high overhead was a family of gibbons moving through the distant branches. Certainly it was a climactic moment, but even though I felt a burst of exhilaration, it was quickly followed by a sense of alarm. I was shocked by how hard it was to see the animals, which compared to their surroundings looked more like squirrels than apes, and I began to wonder how I was ever going to follow them for a full day, let alone simultaneously collect behavioral data. An even bigger surprise, as I later wrote in my diary, was how difficult it was to see the fruit the gibbons had apparently come to the tree to eat. I suppose I had expected fruit the size of supermarket produce dangling from every branch. Instead I could barely distinguish the nickel and dime-sized fruit from the branches themselves. This predicament was no small matter because the topic I had come to Khao Yai to study was the feeding behavior of these highly frugivorous primates.
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